Publications
Pennsylvania Natural Heritage Program puts together reports for its completed projects, fact sheets about species and natural communities, lists of special-concern plants and animals in Pennsylvania, and other documents that span the breadth of program activities.
Scientific Publications
Note: The names of PNHP staff are capitalized.
Yawn ND, Determann RO, Folkerts DR, Thompson P, Stephens JD. 2023. Census and reassessment of the critically endangered Alabama Canebrake Pitcher Plant (sarracenia alabamensis), 25 years later 1. The Journal of the Torrey Botanical Society 150(3), 365-384
Shaw A, Duffy A, Nieto-Lugilde M, Aguero B, SCHUETTE S, Robinson S, Loveland J, Hicks K, Weston D, Piatkowski B, Kolton M, Koska J, Healey, Adam. 2023. Clonality, local population structure, and gametophyte sex ratios in cryptic species of the Sphagnum magellanicum complex. Annals of botany. 10.1093/aob/mcad077.
Estes D, Tracey C, ZIMMERMAN E, Knapp W, Vanderhorst J, Singhurst J, Witsell T. 2023. "Riverscour Ecosystems of Eastern Unglaciated North America: A Review," Natural Areas Journal, 43(3), 148-168
Roberts HP, Willey LL, Jones MT, King DI, . . . , GIPE KD. 2023. Effects of landscape structure and land use on turtle communities across the eastern United States. Biological Conservation, 283. https://doi.org/10.1016/j.biocon.2023.110088 .
Roberts, HP, Willey LL, Jones MT, Akre TSB, . . ., GIPE KD, et al. 2023. Is the future female for turtles? Climate change and wetland configuration predict sex ratios of a freshwater species. Global Change Biology. https://doi.org/10.1111/gcb.16625.
Braund J and Sahli H. 2022. Mating system, pollination, and seed dispersal in the Pennsylvania endangered Iris verna. Journal of the Torrey Botanical Society 149(1): 8–16, 2022.
Bowden R, Caylor A, Hemmelgarn G, KRESSE M, Martin A, and Althouse M. 2022. "Prescribed Browsing by Goats Shows Promise in Controlling Multiflora Rose in a Deciduous Forest at the Erie National Wildlife Refuge in Northwestern Pennsylvania," Natural Areas Journal, 42(3), 196-205
CIAFRE CM, Gienger CM, Rehm EM, Estes LD. 2022. Deterministic and stochastic factors jointly drive plant community composition and diversity in isolated wetlands. Wetlands 42:71.
CIAFRE C, Naczi R. 2022. Rhynchospora stiletto (Cyperaceae), a new species of beaksedge from the southeastern U.S.A.. Kew Bulletin. 7. 10.1007/s12225-022-10044-1.
Willey LL, Jones MT, Sievert PR, Akre TSB, . . ., GIPE KD, et al. 2022. Distribution models combined with standardized surveys reveal widespread habitat loss in a threatened turtle species. Biological Conservation 266:e109437. https://doi.org/10.1016/j.biocon.2021.109437
CIAFRE, C, Estes D, Dwayne L. 2022. Two Coastal Plain Dichanthelium (Poaceae: Paniceae) disjunct in Tennessee grasslands and their conservation. Castanea, 87(1), 121-128. 10.2179/0008-7475.87.1.121.
GRUND SP in press 2022. FABACEAE: DESMODIUM: The Identity of Desmodium dillenii Darl. in Weakley AS, D.B. Poindexter DB, Sorrie BA, Ungberg EA, Ward SG, Horn JW, Knapp W, Grund SP. Studies in the Vascular Flora of the Southeastern United States. VIII. J. Bot. Res. Inst. Texas 16(2):377—418.
Eiseman CS, Smith DR, WOODS P. in press 2022. Nearctic "slug" sawfly larvae of the genus Caliroa Costa (Hymenoptera: Tenthrinidae): New rearing records and a summary of hosts, descriptions, and distribution records. Journal of the Proceedings of the Entomological Society of Washington.
Allison Jr PF, LIEB DA, Loughman ZJ. 2022. Distribution, natural history, and conservation of Cambarusdubius in Pennsylvania, Journal of Natural History, 56,13-16, 829-848, the article.
Hamilton H, Smyth RL, Young BE, Howard TG, TRACEY C, Breyer S, Cameron DR, et al. 2022. Increasing Taxonomic Diversity and Spatial Resolution Clarifies Opportunities for Protecting US Imperiled Species. Ecological Applications 32(3), e2534. View Abstract
Noss RF, Cartwright JM, Estes D, Witsell T, Elliott G, Adams D, Albrecht M, Boyles R, Comer P, Doffitt C, Faber-Langendoen D, Hill J, Hunter WC, Knapp WM, Marshall ME, Singhurst J, TRACEY C, Walck J, Weakley A. 2021. Science needs of southeastern grassland species of conservation concern—A framework for species status assessments: U.S. Geological Survey Open-File Report 2021–1047, 58 p. View Abstract
Noss RF, Cartwright JM, Estes D, Witsell T, Elliott G, Adams D, Albrecht M, Boyles R, Comer P, Doffitt C, Faber-Langendoen D, Hill J, Hunter WC, Knapp WM, Marshall ME, Singhurst J, TRACEY C, Walck J, Weakley A. 2021. Improving species status assessments under the U.S. Endangered Species Act and implications for multispecies conservation challenges worldwide. Conserv Biol.35(6), 1715-1724. View the article.
JEWITT A, Antolos E, Lutz C, Dean J. 2021. Targeted species projects for volunteers to increase early detection capacity: The water chestnut mapping challenge. Natural Areas J. 41, 203–8. Click here to view on the BioOne website.
Jones, MT, Willey LL, eds. (GIPE KD, contributor) 2021. Biology and Conservation of the Wood Turtle. Northeast Association of Fish and Wildlife Agencies.
Roberts HP, Jones MT, Willey LL, Akre TSB, Sievert PR, deMaynadier P, Gipe KD. 2021. Large-scale collaboration reveals landscape-level effects of land-use on turtle demography. Global Ecology and Conservation 30:e01759. https://doi.org/10.1016/j.gecco.2021.e01759.
Allison PF Jr., Khan TN, Davis DA, LIEB DA, Loughman ZJ. 2021. Review of the distribution and new occurrences of Lacunicambarus thomai (Jezerinac, 1993) (Decapoda, Cambaridae) in Pennsylvania and its possible introduction outside of its native range. Check List 17(6), 1461-1473. View the article.
Rocco JM, Regan KM, Larkin JL, EICHELBERGER C, Wisgo J, Nealen PM, et al. 2020. Higher prevalence of Babesia microti than Borrelia burgdorferi in small mammal species in central Pennsylvania, United States. Vector Borne Zoonotic Diseases. 20, 151–154. doi: 10.1089/vbz.2019.2493
SCHUETTE S, Folk RA, Cantley JT, Martine CT. 2018. The hidden Heuchera: How science Twitter uncovered a globally imperiled species in Pennsylvania, USA. PhytoKeys:87–97.
Loughman ZJ, WOODS P and LIEB DA 2018. First Record of Creaserinus fodiens (Digger Crayfish) from Pennsylvania, Northeastern Naturalist, 25(2), N15-N18. View the article.
Glon MG, Mularo AJ, LIEB DA, and Loughman ZJ. 2018. Rediscovery of Cambarus diogenes (Devil Crayfish) in Pennsylvania, Northeastern Naturalist 25(3):355-361. View the article.
SCHUETTE S and TRACEY C. 2017. An Updated List of Bryophytes from Erie County, Pennsylvania, Evansia 34(2):40-53, (1 June 2017). View Abstract
Loughman ZJ, LIEB DA, Scott RK, Dillard ZW, Sadecky NM, 2017. Historical and current distribution of Appalachian primary burrowing crayfishes (Decapoda: Astacoidea: Cambaridae) in western Pennsylvania: a century of change or stasis?, Journal of Crustacean Biology 37:535–543. View the article.
Leuenberger W, Bearer S, Duchamp J, Johnson S, LEPPO B, McElhenny P, Larkin J. 2016. A Comparison of Lepidoptera Communities Inhabiting Restored and Late Successional Pitch Pine—Scrub Oak Barrens in Pennsylvania. Natural Areas Journal 36:38–47.
Ganger MT, ZIMMERMAN EA, GRUND SP, Bissell JK. 2016. The vascular plant flora and plant communities of Erie Bluffs State Park, Erie County, Pennsylvania. Rhodora 118:148–188.
Shiffer CN, Street SG, LEPPO B, White HB. 2015. Odonata of Beaver Dam, Huntingdon County, Pennsylvania: A Record of Faunal Succession in a Changing Habitat. Argia 27:12.
Zanatta DT et al. 2015. Distribution of Native Mussel (Unionidae) Assemblages in Coastal Areas of Lake Erie, Lake St. Clair, and Connecting Channels, Twenty-Five Years After a Dreissenid Invasion. Northeastern Naturalist 22:223–235.
SCHUETTE S, Krayesky DM. 2014. A List of Bryophytes for Wayne County, Pennsylvania. Evansia 31:25–30.
Jongejans E, Skarpaas O, Ferrari MJ, Long ES, Dauer JT, Schwarz CM, Rauschert ESJ, Jabbour R, Mortensen DA, Isard SA, LIEB DA, Sezen Z, Hulting AG and Shea K. A unifying gravity framework for dispersal. 2014. Theoretical Ecology 8, 207–223. View the article.
Shiffer CN, LEPPO B, White HB. 2014. Odonata of Black Moshannon State Park, Centre County, Pennsylvania. Argia 26:9.
Carline RF and LIEB DA. 2014. Influencing Your Agency's Thinking. Pages 251-255 in Taylor, W.W., A.J. Lynch, and N. Leonard, editors. Future of Fisheries: Perspectives for Emerging Professionals. American Fisheries Society. Bethesda, Maryland.
Vanderhorst JP, Streets BP, Faulkner PL, GRUND SP, KUNSMAN JR. 2013. West Virginia. Castanea 78:134–136.
TRACEY C and WOODS PG, 2012. A New Native Plant for Pennsylvania, Equisetum scirpoides Michx.(Equisetaceae). Rhodora, 114(960):406-408.
LIEB DA, Bouchard RW, Carline RF, Nuttall TR, Wallace JR and Burkholder CL. 2011. Conservation and Management of Crayfishes: Lessons from Pennsylvania, Fisheries, 36:10, 489-507. View the article.
Filipova L, LIEB DA, Grandjean F, Petrusek A. 2011. Haplotype variation in the spiny-cheek crayfish Orconectes limosus: colonization of Europe and genetic diversity of native stocks. J. N. Am. Benthol. Soc., 2011, 30(4):871–881. View the article.
LIEB DA, Bouchard RW, Carline RF. 2011. Crayfish Fauna of Southeastern Pennsylvania: Distributions, Ecology, and Changes Over the Last Century, Journal of Crustacean Biology, 31(1):166-178. View the article.
EVANS RR and Ray SJ. 2010. Distribution and environmental influences on freshwater gastropods from lotic systems and springs in Pennsylvania, USA, with conservation implications. American Malacological Bulletin 28:135-150. Click here to view the abstract on the BioOne website.
Smith TA and Crabtree D. 2010. Freshwater Mussel (Unionidae: Bivalvia) Distributions and Densities in French Creek, Pennsylvania. Northeastern Naturalist 17:387–414.
Smith TA and Meyer ES. 2010. Freshwater Mussel (Bivalvia: Unionidae) Distributions and Habitat Relationships in the Navigational Pools of the Allegheny River, Pennsylvania. Northeastern Naturalist 17:541–564.
Crabtree DL and Smith TA. 2009. Population Attributes of an Endangered Mussel, Epioblasma Torulosa Rangiana (Northern Riffleshell), in French Creek and Implications for Its Recovery. Northeastern Naturalist 16:339–354.
HART JA, Wampler T, Krutzsch P. 2009. The Seminole Bat (Lasiurus seminolus) and Virginia Big-eared Bat (Corynorhinus townsendii virginianus) in Pennsylvania. Bat Research News 50(1):1-4.
Chapman EJ and SMITH TA. 2008. Structural community changes in freshwater mussel populations of Little Mahoning Creek, Pennsylvania. American Malacological Bulletin 26:161-169. Click here to view the abstract on the BioOne website.
EVANS R and RAY S. 2008. Checklist of the freshwater snails (Mollusca: Gastropoda) of Pennsylvania, USA. Journal of the Pennsylvania Academy of Science 82(2/3):92–97.
Ferster B, LEPPO BR, Swartz MT, Vulinec K, Habegger F, Mehring A. 2008. Lepidoptera of Fort Indiantown Gap National Guard Training Center, Annville, Pennsylvania. Northeastern Naturalist 15(1):141–148. Click here to view the abstract on the BioOne website.
LIEB DA, Carline RF, Rosenberger JL, and Mengel VM. 2008. The discovery and ecology of a member of the Cambarus acuminatus complex (Decapoda: Cambaridae) in Valley Creek, southeastern Pennsylvania. Journal of Crustacean Biology 28: 439-450.
Pearce TA and EVANS R. 2008. Freshwater Mollusca of Plummers Island, Maryland. Bulletin of the Biological Society of Washington 15(1): 20–30. Click here to view abstract on the BioOne website.
GRUND SP and Isaac BL. 2007. Taxonomy and Lectotypification of Appalachian Blue Violet. Castanea 72:58–61.
JELLEN BC and KOWALSKI MJ. 2007. Movement and growth of neonate Eastern Massasaugas
(Sistrius catenatus). Copeia 2007(4):994–1000. Click
here to view the abstract on the BioOne website.
GRUND SP and Parks JC. 2003. Conservation status of pteridophytes in the 49 continental United States: A preliminary report. Fern Gaz. 16(6): 290-294.
Reports
CIAFRé CM. 2021. Vegetation classification and mapping at Longwood Gardens. Final project report.
YEANY, D. 2020. Evaluating Pennsylvania’s Grasslands to Prioritize Conservation Efforts and Provide the Appropriate Management in the Beneficial Areas for Grassland Nesting Birds. Pennsylvania Natural Heritage Program Final Report for Pennsylvania Game Commission. Cooperative Agreement 4000021689. View abridged version.
Inventories of Pennsylvania's state parks
Natural Resources Inventory and Management Recommendations for Kettle Creek State Park (October 2009).
Natural Resources Inventory and Management Recommendations for Sinnemahoning State Park (June 2008).
Inventory of Rare Plant Species and Plant Communities at Canoe Creek State Park, Blair County, Pennsylvania: Progress Report (June 2006).
Inventory of Rare Species and Plant Communities at Black Jack Swamp Natural Area and Clark Island Natural Area, Pymatuning State Park, Crawford County, Pennsylvania: Final Report (September 2004).
Inventory of Rare Species and Plant Communities, Slippery Rock Natural Area, Moraine State Park, Butler County, Pennsylvania: Final Report (April 2004).
Inventory of Rare Species and Plant Communities at Forest Cathedral Natural Area and Swamp Forest Natural Area, Cook Forest State Park, Clarion and Forest Counties, Pennsylvania: Final Report (August 2003).
Inventory of Rare Species and Plant Communities at Hemlock Trail Natural Area, Laurel Hill State Park, Somerset County, Pennsylvania: Final Report (August 2003).
Inventory of Rare Species and Plant Communities, Black Moshannon Bog Natural Area, Black Moshannon State Park, Centre County, Pennsylvania: Final Report (July 2003).
Inventories and vegetation classification of National Park Service units in Pennsylvania
Classification and Mapping of Vegetation and Fire Fuel Models at Delaware Water Gap National Recreation Area (2007). Click here to access the report on the National Park Service website.
Vegetation Classification and Mapping at Allegheny Portage Railroad National Historic Site (2007). Click here to access the report on the National Park Service website.
Vegetation Classification and Mapping at Gettysburg National Military Park and Eisenhower National Historic Site (2006). Click here to access the report on the National Park Service website.
Vegetation Classification and Mapping at Friendship Hill National Historic Site (2006). Click here to access the report on the National Park Service website.
Vegetation Classification and Mapping at Fort Necessity National Battlefield (2006). Click here to access the report on the National Park Service website.
Vegetation Classification and Mapping at Johnstown Flood National Memorial (2006). Click here to access the report on the National Park Service website.
Inventory of Plant Species of Special Concern at Gettysburg National Military Park and Eisenhower National Historic Site (2006)
Inventory of Bat Community Composition at Valley Forge National Historical Park (2006). Click here to access a pdf of the report on the National Park Service website.
Inventory of Bat Community Composition at Hopewell Furnace National Historic Site. (2006). Click here to accesss a pdf of the report on the National Park Service website.
Inventory of Bat Community Composition at Gettysburg National Military Park and Eisenhower National Historic Site (2006). Click here to access a pdf of the report on the National Park Service website.
Survey for Endangered and Threatened Mammals at Gettysburg National Military Park and the Eisenhower National Historic Site (2006)
Vegetation Classification and Mapping of Valley Forge National Historical Park (2005). Click here to access the report on the National Park Service website.
Vegetation Classification and Mapping of Hopewell Furnace National Historic Site (2005). Click here to access the report on the National Park Service website.
Community Classifications
Terrestrial and Palustrine Plant Communities of Pennsylvania, 2nd Edition (2012).
Pennsylvania Statewide Seasonal Pool Ecosystem Classification (2009).
Classifying Lotic Systems for Conservation: Methods and Results of the Pennsylvania Aquatic Community Classification (June 2007). Click here to view.
User's Manual and Data Guide to the Pennsylvania Aquatic Community Classification (June 2007). Click here to view.
Terrestrial and palustrine plant communities of Pennsylvania. Pennsylvania Natural Diversity Inventory. Pennsylvania Department of Conservation and Recreation. (June 1999)
A Study of Seepage Wetlands in Pennsylvania (March 1998).
A Study of Calcareous Fen Communities in Pennsylvania (1995).
